Mark Wilkinson - Publications 

Undescribed Siphonops

PDFs of many of these are available upon request


2009| 2008| 2007| 2006| 2005| 2004| 2003| 2002| 2001| 2000| 1999| 1998| 1997| 1996| 1995| 1994| up to 1992|


Doherty-Bone, T. M., Kebuh Ndifon, R., San Mauro, D., Wilkinson, M., Nono LeGrand, G. N. & Gower, D. J. 2011. Systematics and ecology of the caecilian Crotaphatrema lamottei (Nussbaum) (Amphibia: Gymnophiona: Scolecomorphidae). Journal of Natural History 45:827-841.

Giri, V., Gower, D. J., Gaikwad, K. & Wilkinson, M. 2011. A second species of Gegeneophis Peters (Amphibia: Gymnophiona: Caeciliidae) lacking secondary annular grooves. Zootaxa 2815:49-58.

Gower, D. J., ,Papadopoulou, A., Doherty-bone, T. M., Pupin, F., San Mauro, D. Loader, S. P. & Wilkinson, M. 2011. The systematics of Boulengerula fischeri (Amphibia: Gymnophiona: Caeciliidae) based on morphological and molecular data. Zootaxa 2767:14-24.

Gower, D. J., San Mauro, D., Giri, V., Bhatta, G., Govindappa, V., Kotharambath, R., Oommen, O. V., Fatih, F. A., Mackenzie-Dodds, J. A., Nussbaum, R. A., Biju, S. D., Shouche, Y. S. & Wilkinson, M. 2011. Molecular systematics of caeciliid caecilians (Amphibia: Gymnophiona) of the Western Ghats, India. Molecular Phylogenetics and Evolution 59:698-707.

Loader, S. P., Wilkinson, M., Cotton, J. A., Measey, G. J., Menegon, M., Howell, K. H., MŸller, H. & Gower, D. J. 2011. Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. The Herpetological Journal 21:5-16.

McMorris, F. R. & Wilkinson, M. 2011. Conservative supertrees. Systematic Biology 60:232-238.

Wilkinson, M. 2011. Compatibility analysis and data quality in morphological and molecular phylogenetics. P. 390. In (Borsch, T., Giere, P., Hoffman, J., Jahn, R., Lšhne, C,, Nordt, B. & Ohl, M. eds.) Biosystematics Berlin 2011, Programme and Abstracts..

Wilkinson, M. & Day, J. J. 2011. Testing hard polytomies with partial splits. Zitelliana 30:56.

Wilkinson, M., San Mauro, D., Sherratt, E, & Gower, D. J. 2011. A nine-family classification of caecilian amphibians. Zootaxa 2874:41-64.


Gower, D. J., Wilkinson, M., Sherratt, E. & Kok, P. J. R. 2010. A new species of Rhinatrema (Amphibia: Gymnophiona: Rhinatrematidae) from Guyana. Zootaxa 2391:47-60.

Hoffman, M et al. 2010. The impact of conservation on the status of the WorldÕs vertebrates. Science 330:1503-1509.

Houston, A. I., Wood, J. & Wilkinson, M. 2010. How did the Great Auk raise its young? Journal of Evolutionary Biology 23:1899-1906.

Mohun, S. M., Davies, W. L., Bowmaker, J. M., Pisani, D., Himstedt, W., Gower, D. J., Hunt, D. M. & Wilkinson, M. 2010. Identification and characterization of visual pigments in caecilians, a group of limbless amphibians with rudimentary eyes that form the Order Gymnophiona.. Journal of Experimental Biology 213:3586-3592.

Wilkinson, M. & Gower, D. J. 2010. A new species of Rhinatrema (Amphibia: Gymnophiona: Rhinatrematidae) from Amazonas, Brazil. Zootaxa 2650:63-68.

Wilkinson, M. & Kok, P. J. R. 2010. A new species of Microcaecilia (Amphibia: Gymnophiona: Caeciliidae) from Guyana. Zootaxa 2719:35-40.


Baker, W. J., Savolainen V., Asmussen-Lange, C. B., Chase, M. W., Dransfield, J., Forest, F., Harley, M.H., Uhl, N. W. & Wilkinson, M. 2009. Complete generic level phylogenetic analyses of palms (Arecaceae) with comparisons of supertree and supermatrix approaches. Systematic Biology 58:240-256.

Gower, D. J., Sherratt, E. & Wilkinson, M. 2009. A pilot study of the systematics and ecology of caecilian amphibians of the Station de Recherche des Nouragues. Report for CNRS. Pp. 1-15.

Gower, D. J. & Wilkinson, M. 2009. Gymnophiona. Pp. 369-372. In (S. B. Hedges & S. Kumar, eds.) The Timetree of Life. Oxford University Press, New York.

Cotton, J. A. & Wilkinson, M. 2009. Supertrees enter the mainstream of phylogenetics. Trends in Ecology and Evolution 24:1-3. 

Müller, H., Wilkinson, M., Loader, S. P., Wirkner, C. S. & Gower, D. J. 2009. Morphology and function of the head in foetal and juvenile Scolecomorphus kirkii (Amphibia: Gymnophiona: Scolecomorphidae). Biological Journal of the Linnean Society 96:491-504.

Kamei, R. G., Wilkinson, M., Gower, D. J. & Biju, S. D. 2009. Three new species of striped Ichthyophis (Amphibia: Gymnophiona: Ichthyophidae) from the northeast Indian states of Manipur and Nagaland. Zootaxa 2267:26-42.

Loader, S. P., Gower, D. J., Wilkinson, M., Menegon, M. & Müller, H. 2009. The endemic amphibians of the Nguru Mountains. Declining Amphibian Populations Taskforce Report. Pp 1-22.

Loader, S. P., Menegon, M., Müller, H., Gower, D. J., Wilkinson, M., Howell, K., & Orton, F. 2009. Remarkable amphibian diversity in the South Nguru Mountains, Tanzania. Froglog 91:8-10.

San Mauro, D., Gower, D. J., Massingham, T., Wilkinson, M., Zardoya, R. & Cotton, J. A. 2009. Experimental design in caecilian systematics: phylogenetic information of mitochondrial genomes and nuclear rag1. Systematic Biology 58:425-438.

Wilkinson, M., Hoogmoed, M. S. & Nussbaum, R. A. 2009. A new species of Microcaecilia (Amphibia: Gymnophiona: Caeciliidae) from Suriname. Herpetologica 65:413-418.


Bain, R., Biju, S.D., Brown, R., Das, I., Diesmos, A., Dutta, S., Gower, D., Inger, R., Iskandar, D., Kaneko, Y., Lau, M.W.N., Meegaskumbura, M., Ohler, A., Papenfuss, T., Pethiyagoda, R., Stuart, B., Wilkinson, M. & Xie, F. 2008. Amphibians of the Indomalayan region. Pp.74-79. In (S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani & B. E. Young, eds.) Threatened Amphibians of the World. Lynx Ediciones, with IUCN - The World Conservation Union, Conservation International, and NatureServe, Barcelona.

Bola–os, F., Castro, F., Cortez, C., De la Riva, I., Grant, T., Hedges, S. B., Heyer, R., Ib‡–ez, R., La Marca, E., Lavilla, E., Leite SIlvano, D., Lštters, S., Para Olea, G., Reiche, S., Reynolds, R., Rodr’guez, L.., Santos Barrera, G., Scott, N., Ubeda, C., Veloso, A., Wilkinson, M. & Young, B. 2008. Amphibians of the Neotropical Realm. Pp.95-105. In (S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani & B. E. Young, eds.) Threatened Amphibians of the World. Lynx Ediciones, with IUCN - The World Conservation Union, Conservation International, and NatureServe, Barcelona.

Cotton, J. A. & Wilkinson, M. 2008. Quantifying the potential utility of phylogenetic characters. Taxon 57:131-136.

Day, W., McMorris, F. R. & Wilkinson, M. 2008. Explosions and hotspots in supertree methods. Journal of Theoretical Biology 253:345-348.

Du Preez, L. H., Wilkinson, M. & Huyse, T. 2008. The first record of polystomes (Monogenea: Polystomatidae) from caecilian hosts (Amphibia: Gymnophiona) with the description of a new genus and two new species. Systematic Parasitology 69:201-209.

Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., de Sa;. R. O., Channing, A., Wilkinson, M., Donnellan, S. C. Raxworthy, C. J., Campbell, J. A., Blotto, B., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J., Green, D. M. & Wheeler, W. C. 2008. Is the amphiban tree of life really fatally flawed? Cladistics 24:385-395.

Gower, D. J., Rajendran, A., Nussbaum, R. A. & Wilkinson, M. 2008. A new species of Uraeotyphlus (Amphibia: Gymnophiona: Uraeotyphlidae) of the malabaricus group. Herpetologica 64:235-245.

Gower, D. J. & Wilkinson, M. 2008. Caecilians (Gymnophiona). Pp. 19-20. In (S. N. Stuart, M. Hoffmann, J. S. Chanson, N. A. Cox, R. Berridge, P. Ramani & B. E. Young, eds.) Threatened Amphibians of the World. Lynx Ediciones, with IUCN - The World Conservation Union, Conservation International, and NatureServe, Barcelona.

Kupfer, A., Wilkinson, M., Gower, D. J., Müller, H. & Jehle, R. 2008. Care and parentage in a skin-feeding caecilian amphibian. Journal of Experimental Zoology 309A:460-467.

Wilkinson, M., Kupfer, A., Marques-Porto, R., Jeffkins, H., Antoniazzi, M. M., Jared, C. 2008. One hundred million years of skin feeding? Extended parental care in a Neotropical caecilian (Amphibia: Gymnophiona). Biology Letters 4:358-361.


Cobbett, A., Wilkinson, M. & Wills, M. A. 2007. Fossils impact as hard as recent taxa in parsimony analysis. Systematic Biology. 56:753-766.

Cotton, J. A. & Wilkinson, M. 2007. Majority rule supertrees.   Systematic Biology 56:445-452.

Grazziotin F. G., Zaher, H., Ferrarezzi, H., Klaczko J., Bonato, S. L. & Wilkinson, M. 2007. Higher-level molecular phylogeny of snakes: conflicts and congruence. Darwiniana supplement 45:17-19.

Gower, D. J., Dharne, M., Bhatta, G. Giri, V. Vyas, R., Govindappa, V., Oommen, O. V., George, J, Schouche Y. & Wilkinson, M. 2007. Remarkable genetic homogeneity in unstriped, long-tailed Ichthyophis (Amphibia: Gymnophiona: Ichthyophiidae)along 1500 km of the Western Ghats, India. Journal of Zoology. 272:266-275.

Gower, D. J., Giri, V. & Wilkinson, M. 2007. Rediscovery of Gegeneophis seshachari Ravichandran, Gower & Wilkinson, 2003 at the type locality. Herpetozoa 19:121-127.

Gower, D. J.. & Wilkinson, M. 2007. Species groups in Uraeotyphlus (Amphibia: Gymnophiona: Uraeotyphiidae) with the description of a new species. Herpetologica 63:401-410.

Harris, S. R., Pisani, D., Gower, D. J. & Wilkinson, M. 2007. Investigating stagnation in morphological phylogenetics using consensus data. Systematic Biology 56:125-129.

Loader, S. P., Pisani, D., Cotton, J. A., Gower, D. J., Day, J. J. & Wilkinson, M. 2007. Relative timescales reveal multiple origins of parallel disjunct distributions of African caecilian amphibians. Biology Letters. 3:505-508.

Pisani, D., Benton, M. J. & Wilkinson, M. 2007. Congruence of morphological and molecular phylogenies. Acta Biotheoretica 55:269-281.

Roelants, K., Gower, D. J., Wilkinson, M., Loader, S. P., Biju, S. D., Guillaume, K. & Bossuyt, F. 2007. Patterns of diversification in the history of modern amphibians. Proceedings of the National Academy of Sciences. 104:887-892.

Wilkinson, M., Cotton, J. A., Lapointe, F.-J. & Pisani, D. 2007. Properties of supertree methods in the consensus setting. Systematic Biology 56:330-337.

Wilkinson, M., Gower, D. J., Govindappa, V. & Venkatachalaiah. 2007. A new speces of Ichthyophis (Amphibia: Gymnophiona: Ichthyophiidae) from Karnataka, India. Herpetologica 63:511-518.

Wilkinson, M., McInerney, J. O., Hirt, R. P., Foster, P. & Embley, T. M. 2007. Of clades and clans: terms for relationships in unrooted trees. Trends in Evolution and Ecology 22:114-115.


Cotton, J. A., Slater, C. & Wilkinson, M. 2006. Discriminating unsupported and supported relationships in supertrees using triplets. Systematic Biology 55:345-350.

Day, J. J. & Wilkinson, M. 2006. On the origin of the Synodontis catfish species flock from Lake Tanganyika.<Biology Letters 2:548-552.

Frost, D. R., Grant, T., Faivovich, J., Bain, R. H., Haas, A., Haddad, C. F. B., de Sa, R. O., Channing, A., Wilkinson, M., Donnellan, S. C. Raxworthy, C. J., Campbell, J. A., Blotto, B., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J., Green, D. M. & Wheeler, W. C. 2006. The amphiban tree of life. Bulletin of the American Museum of Natural History 297:1-370.

Gower, D. J., Oommen, O. V. & Wilkinson, M. 2006. Marking amphibians with alpha numeric fluorescent tags - caecilians lead the way. Herpetological Review 37:302.

Kupfer, A., Gower, D. J., Gaucher, P. & Wilkinson, M. 2006. Passive trapping of aquatic caecilians (Amphibia: Gymnophiona: Typhlonectidae). Studies on Neotropical Fauna and Environment 41: 93-96.

Kupfer, A., Müller, H., Jared, C., Antoniazzi, M., Nussbaum, R. A., Greven, H.& Wilkinson, M. 2006. Parental investment by skin feeding in a caecilian amphibian. Nature 440:926-929.

Loader, S. P. & Wilkinson, M. 2006. Draft report on the reptiles and amphibians of the Bumbuna region, Sierra Leonne. Confidential Report for the World Bank. 45 pp.

Pisani, D., Mohun, S. M., Harris, S. R., McInerney, J. O. & Wilkinson, M. 2006. Molecular evidence for dim-light vision in the last common ancestor ofthe vertebrates. Current Biology 16:R318-319.

San Mauro, D., Gower, D. J., Zardoya, R. & Wilkinson, M. 2006. A hotspot of gene order rearrangement by tandem duplication and random loss in the vertebrate mitochondrial genome. Molecular Biology and Evolution 23:227-234.

Venkatachalaiah, G., Govindappa, V., Giri, V. & Wilkinson, M. 2006. Studies on the karyotype of Indotyphlus battersbyi Taylor (Amphibia: Gymnophiona: Caeciliidae). The Journal of Cytology and Genetics 7:47-56.

Wilkinson, M. 2006. Identifying stable reference taxa for phylogenetic nomenclature. Zoologica Scripta 35:109-112.

Wilkinson, M. & Cotton, J. A. 2006. Supertree methods for building the tree of life: divide-and-conquer approaches to large phylogenetic problems. In T. R. Hodkinson & J. A. N. Parnell (Eds.) Reconstructing the Tree of Life: Taxonomy and Systematics of Large and Species Rich Taxa. Systematics Association Special Volume 72. CRC Press.

Wilkinson, M. & Nussbaum, R. A. 2006. Caecilian phylogeny and classification. In J.-M. Exbrayat (Ed.) Reproductive Biology and Phylogeny of Amphibia, Volume 3. Gymnophiona. Science Publishers Inc. Pp. 39-78.


Bossuyt, F., Meegaskambura, M. Beenaerts, N., Gower, D. J., Pethiyagoda, R., Roelants, K., Mannaert, A., Wilkinson, M., Bahir, M. M., Manamendra-Arachchi, K., Ng, P. K. L., Schnieider, C. J., Oommen O. V. & Milinkovitch, M. C. 2005. Biodiversity in Sri Lanka and the Western Ghats: Response. Science 308:199.

Gower D. J., & Wilkinson, M. 2005. The conservation biology of caecilians. Conservation Biology 19:45-55.

Gower, D. J. Bahir, M. M., Mapatuna, Y., Pethiyagoda, R., Raheem, D. & Wilkinson, M. 2005. Molecular phylogenetics of Sri Lankan Ichthyophis (Amphibia: Gymnopohiona: Ichthyophiidae), with discovery of a cryptic species. Raffles Bulletin of Zoology, Supplement No. 12:249-257.

McInerney, J. O. & Wilkinson, M. 2005. New methods ring changes for tree of life. Trends in Ecology and Evolution 20:105-107.

Wilkinson, M., Pisani, D., Cotton, J. A. & Corfe, I. 2005. Measuring support and finding unsupported groups in supertrees. Systematic Biology 54:823-831.

Wilkinson, M., Cotton, J. A., Creevey, C., Eulenstein, O., Harris, S. R., Lapointe, F.-J., Levasseur, C., McInereny J. O., Pisani, D. & Thorley, J. L. 2005. The shape of supertrees to come. tree shape related properties of fourteen supertree methods. Systematic Biology 54:419-431.


Bossuyt, F., Meegaskambura, M. Beenaerts, N., Gower, D. J., Pethiyagoda, R., Roelants, K., Mannaert, A., Wilkinson, M., Bahir, M. M., Manamendra-Arachchi, K., Ng, P. K. L., Schnieider, C. J., Oommen O. V. & Milinkovitch, M. C. 2004. Local endemism within the Western Ghats-Sri Lanka biodiversity hotspot. Science 306:471-489.

Creevey, C. J., Fitzpatrick, D. A., Philip, G. K, Kinsella, R. J., O'Connell, M. J., Pentony, M. M., Travers, S. A., Wilkinson, M. & McInerney, J. O. 2004. Does a tree-like phylogeny only exist at the tips in the prokaryotes? Proceedings of the Royal Society B 271:2551-2558.

Giri, V., Gower, D. J. & Wilkinson, M. 2004. A new species of Indotyphlus Taylor (Amphibia: Gymnophiona: Caeciliidae) from the Western Ghats, India. Zootaxa 739:1-19.

Gower, D. J., Bhatta, G., Giri, V. Oommen, O. V., Ravichandran, M. S. & Wilkinson, M. 2004. Biodiversity in the Western Ghats: the discovery of new species of caecilian amphibians. Current Science 87:739-740.

Gower, D. J., Loader, S. P. & Wilkinson, M. 2004. Assessing the conservation status of soil-dwelling vertebrates: insights from the rediscovery of Typhlops uluguruensis (Reptilia: Serpentes: Typhlopidae). Systematics and Biodiversity 2:79-82.

Gower, D. J., Loader, S. P., Wilkinson, M. & Moncreiff, C. B. 2004. Niche separation and comparative abundance of Boulengerula boulengeri and Scolecomorphus vittatus (Amphibia: Gymnophiona) in East Usambara forest, Tanzania. African Journal of Herpetology 53:183-190.

Gower, D. J., Rasmussen, J. B., Loader, S. P. & Wilkinson, M. 2004. The caecilian amphibian Scolecomorphus kirkii Boulenger as prey of the burrowing asp Atractaspis aterrima GŸnther: Trophic relationships of fossorial vertebrates. African Journal of Ecology, 42:83-87.

Loader, S. P., Gower, D. J., Howell, K. M., Doggart, N., Rödel, M.-O., Clarke, B. T., de Sa, R. O., Cohen, B. L. & Wilkinson, M. 2004. Phylogenetic relationships of African microhylid frogs inferred from DNA sequences of mitochondrial 12s and 16s rRNA genes. Organisms, Diversity and Evolution 4:227-235.

Measey, G. J., Gower, D. J. ,Oommen, O. V. & Wilkinson, M. 2004. A subterranean generalist predator: diet of the fossorial caecilian Gegeneophis ramaswamii (Amphibia; Gymnophiona; Caeciliidae) in southern India. Compte Rendus Biologies 327:65-76.

Møbjerg, N, Jespersen, Å & Wilkinson, M. 2004. Morphology of the kidney in the West African caecilian, Geotrypetes seraphini (Amphibia, Gymnophiona, Caeciliidae). Journal of Morphology 262:583-607.

San Mauro, D., Gower, D. J., Oommen O.V., Wilkinson, M. & Zardoya, R. 2004. Phylogeny of caecilian amphibians (Gymnophiona) based on complete mitochondrial genomes and nuclear RAG1. Molecular Phylogenetics and Evolution 33:413-427.

Wilkinson, M., Cotton, J. A. & Thorley, J. L. 2004. The information content of trees and their matrix representations. Systematic Biology, 53:989-1001.

Wilkinson, M., Loader, S. P., Müller, H. & Gower, D. J. 2004. Taxonomic status and phylogenetic relationships of Boulengerula denhardti Nieden, 1912 (Amphibia: Gymnophiona: Caeciliidae). Mitteilungen Mus. Natkd. Berlin, Zoologische Reihe, 80:41-51.

Wilkinson, M., Thorley, J. L., Pisani, D. Lapointe, F.-J., and McInerney, J. O. 2004. Some desiderata for liberal supertrees. Pp. 227-246. In (O. Bininda-Emonds, ed.) Phylogenetic Supertrees Kluwer Academic Publishers, Dordrecht, The Netherlands.


Giri, V., Gower, D. J. & Wilkinson, M. 2003. A new species of Gegeneophis Peters (Amphibia: Gymnophiona: Caeciliidae) from southern Maharashtra, India, with a key to the species of the genus. Zootaxa 351:1-10.

Harris, S. R., Gower, D. J. & Wilkinson, M. 2003. Intraorganismal homology, character construction and the phylogeny of aetosaurian archosaurs (Reptilia, Diapsida). Systematic Biology 52:239-252.

Harris, S. R., Gower, D. J. & Wilkinson, M. 2003. Phylogenetic methods and aetosaur interrelationships: a rejoinder. Systematic Biology 52: 851-852.

Harris, S. R., Wilkinson, M. & Marques, A. 2003. Countering concerted homoplasy. Cladistics 19:128-130.

Lapointe, F.-J., Wilkinson, M. & Bryant, D. 2003. Matrix representations with parsimony or with distances: Two sides of the same coin? Systematic Biology, 52:865-868.

Loader, S. P., Gower, D. J. & Wilkinson, M. 2003. Caecilians: mysterious amphibians of the Eastern Arc mountains. Arc Journal 15: 3-4.

Loader S. P., Wilkinson, M., Gower, D. J. & Msuya, C. A. 2003. A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona: Scolecomorphidae) from the North Pare Mountains, Tanzania. Journal of Zoology 259:93-101.

Measey G. J., Gower, D. J., Oommen, O. V., & Wilkinson, M. 2003. Mark recapture study of Gegeneophis ramaswamii (Amphibia: Gymnophiona: Caeciliidae) at a breeding site in Southern Kerala. Journal of Zoology 261:129-133.

Measey, G. J., Gower, D. J., Oommen O. V. & Wilkinson, M. 2003. Quantitative surveying of limbless endogeic soil vertebrates - a case study of Gegeneophis ramaswamii(Amphibia: Gymnophiona: Caeciliidae) in southern India. Applied Soil Ecology 23:43-53.

Pisani, D. & Wilkinson, M. 2003. Software Review, MacClade 4. The Palaeontological Association Newsletter 53: 93-94.

Ravichandran, M. S., Gower, D. J. & Wilkinson, M. 2003. A new species of Gegeneophis Peters (Amphibia: Gymnophiona: Caeciliidae) from Maharashtra, India. Zootaxa 350:1-8.

Scheltinga, D. M., Wilkinson, M., Jameison, B. G. M. & Oommen, O. V. 2003. Ultrastructure of the mature caecilian spermatozoa (Amphibia: Gymnophiona). Journal of Morphology 258:179-192.

Thorley, J. L. & Wilkinson, M. 2003. A view of supertree methods. Pp 185-194. In (M. F. Janovitz, F.-J. Lapointe, F. R. McMorris, B. Mirkin & F. S. Roberts, eds.) Bioconsensus. DIMCAS Series in Discrete Mathematics and Theoretical Computer Science volume 61. The American Mathematical Society.

Wilkinson, M. 2003. Missing data and multiple trees: stability and support. Journal of Vertebrate Paleontology 23:311-323.

Wilkinson, M. Lapointe, F. J. & Gower, D. J. 2003. Branch lengths and support. Systematic Biology 52:127-130.

Wilkinson, M & Loader S P 2003. Worms of wisdom. BBC Wildlife 21:88.

Wilkinson, M., Loader, S. P., Gower, D. J., Sheps, J. A. & Cohen, B. L. 2003. Phylogenetic relationships of African caecilians (Amphibia: Gymnophiona): insights from mitochondrial rRNA gene sequences. African Journal of Herpetology 52:83-92.

Wilkinson, M.,Müller, H. & Gower, D. J. 2003. On Herpele multiplicata (Amphibia: Gymnophiona: Caeciliidae). African Journal of Herpetology 52: 119-122.

Wilkinson, M. & Thorley J. L. 2003. Reduced consensus. Pp. 195-204. In (M. F. Janovitz, F.-J. Lapointe, F. R. McMorris, B. Mirkin & F. S. Roberts, eds.) Bioconsensus. DIMCAS Series in Discrete Mathematics and Theoretical Computer Science volume 61. The American Mathematical Society.


Gower, D. J. & Wilkinson, M. 2002. Phallus morphology in caecilians and its systematic utility. Bulletin of the Natural History Museum (Zoology) 68:143-154.

Gower, D. J., Kupfer, A., Oommen, O. V., Himstedt, W., Nussbaum, R.A., Loader, S. P., Presswell, B., M¨ller, H., Krishna, S. B., Boistel, R. & Wilkinson, M. 2002. A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): Out of India or out of Eurasia? Proceedings of the Royal Society, B 269: 1563-1569.

Knapp, S., Bateman, R. M., Chalmers, N. R., Humphries, C. J., Rainbow, P. S., Smith, A. B., Taylor, P. D, Vane-Wright, R. L. & Wilkinson, M. 2002. Taxonomy needs evolution, not revolution. Nature 419:559.

Pisani, D. & Wilkinson, M. 2002. MRP, Taxonomic Congruence and Total Evidence. Systematic Biology 51:151-155.

Presswell, B., Gower, D. J., Oommen, O. V., Measey, G. J. & Wilkinson, M. 2002. Scolecophidian snakes in the diets of south asian caecilian amphibians. Herpetological Journal 12: 123-126.

Wilkinson, M. 2002. The Narrow Way. Journal of Paleontology 76:1113-1114.

Wilkinson, M., Foster, P. G., Moncrieff, C. B. & Peres-Nuno, P. 2002. Type I error rates of the parsimony permutation tail probability. Systematic Biology 51: 524-527.

Wilkinson, M., Richardson, M. K., Gower, D. J. & Oommen, O. V. 2002. Extended embryo retention, caecilian oviparity and amniote origins. Journal of Natural History 36:2185-2198.

Wilkinson, M., Oommen, O. V., Sheps, J. A. & Cohen, B. L. 2002. Phylogenetic relationships of Indian caecilians (Amphibia: Gymnophiona) inferred from mitochondrial rRNA gene sequences. Molecular Phylogenetics and Evolution 23:401-407.


Harcourt-Brown, K. G., Pearson, P. N. & Wilkinson, M. 2001. The imbalance of paleontological trees. Paleobiology 27: 188-204.

Measey G. J., Gower, D. J. Oommen, O. V. & Wilkinson, M. 2001. Permanent marking of a fossorial caecilian, Gegeneophis ramaswamii (Amphibia: Gymnophiona: Caeciliidae). South Asian Journal of Natural History 5:109-115.

Wilkinson, M. 2001. Phylogenetic Analysis. Pp. 509-515. In (D. E. G. Briggs and P. R. Crowther, eds.) Palaeobiology II. Blackwell. Oxford.

Wilkinson, M. 2001. Cladistics. In:Encyclopedia of the life sciences John Wiley & Sons, Ltd: Chichester, http://www.els.net/ [doi:10.1038/npg.els.0001522]

Wilkinson, M. & Thorley, J. L. 2001. The efficiency of strict consensus trees. Systematic Biology 50:610-613.

Wilkinson, M. & Thorley, J. L. 2001. No compromise on consensus. Taxon 50: 181-184.

Wilkinson, M., Thorley, J. L., Littlewood, D. T. L. & Bray R. A. 2001. Towards a phylogenetic supertree for platyhelminthes? Pp. 292-301 In (D. T L. Littlewood and R. A. Bray, eds) Interrelationships of the Platyhelminthes Taylor & Francis.


Oommen O. V., Measey, G. J., Gower, D. J. & Wilkinson, M. 2000. Distribution and abundance of the caecilian Gegeneophis ramaswamii (Amphibia: Gymnophiona) in southern Kerala. Current Science 79:1386-1389.

Wilkinson, M. 2000. Stability and support: ameliorating the obfuscatory effects of missing data. Journal of Vertebrate Paleontology 20 (3) supplement 79A.

Wilkinson, M., Thorley, J. L. & Upchurch, P. M. 2000. A chain is no stronger than its weakest link: Double decay analyses of phylogenetic hypotheses. Systematic Biology 49:754-776.

Wilkinson, M., Varghese, S. & Oommen, O. V. 2000. Caecilians of Kerala. Biodiversity India 8-12:14.


Thorley, J. L. & Wilkinson, M. 1999. Testing the phylogenetic stability of early tetrapods. Journal of Theoretical Biology 200:343-344.

Viscardi, P., Dyke, G. J., Wilkinson, M. & Rayner, J. M. V. 1999. Missing data and the phylogeny of the Pterosauria.Journal of Vertebrate Paleontology 19 (3) supplement. 82A.

Wilkinson, M. 1999. Choosing and interpreting a tree for the oldest mammal. Journal of Vertebrate Paleontology 19:187-190.

Wilkinson, M. & Nussbaum, R. A. 1999. Evolutionary relationships of the lungless caecilian Atretochoana eiselti (Amphibia: Gymnophiona: Typhlonectidae). Zoological Journal of the Linnean Society 126:91-123.

Zamprogno, C., Zamprogno, M d'G. F. & Wilkinson, M. 1999. Evidence of terrestrial feeding in the "arboreal" lizard Enyalius bilineatus of Eastern Brazil. Bulletin of the British Herpetological Society 67:49-52.


Cohen, B. L., Sheps, J. A. & Wilkinson, M. 1998. Archiving molecular phylogenetic alignments as NEXUS files. Systematic Biology 47: 495-496.

Herniou, E., Martin, J., Miller, K., Cook, J., Wilkinson, M. & Tristem, M. 1998. Retroviral diversity and distribution in the vertebrates. Journal of Virology 72:5955-5966.

Hirt, R. H., Wilkinson, M. & Embley, T. M. 1998. Molecular and cellular evolution of ciliates: a phylogenetic perspective. Pp. 327-340. In (G. H. Coombs, K Vickerman, M. A. Sleigh & A. Warren, Eds.) Evolutionary relationships among protozoa. Kluwer Academic Press.

Measey, G. J. & Wilkinson, M. 1998. Lines of arrested growth in the caecilian Typhlonectes natans (Amphibia: Gymnophiona). Amphibia-Reptilia 19:91-95.

OÕReilly, J. C., Fenolio, D., Rania, L. C. & Wilkinson, M. 1998. Altriciality and extended parental care in the West African caecilian Geotrypetes seraphini (GYMNOPHIONA: CAECILIIDAE).American Zoologist 38: 187A.

Thorley, J. L., Wilkinson, M. & Charleston, M.A. 1998. The information content of consensus trees. Pp. 91-98. In (A. Rizzi, M. Vichi, and H.-H. Bock, eds.) Advances in data science and classification. Springer, Berlin.

Wilkinson, M. 1998. Split support and split conflict randomization tests in phylogenetic inference. Systematic Biology 47:673-695.

Wilkinson, M. 1998. Higher-order homoplasy tests. Acta Biotheoretica 46:109-116.

Wilkinson, M. & Nussbaum, R. A. 1998. Caecilian viviparity and amniote origins. Journal of Natural History 32:1403-1409.

Wilkinson, M. Sebben, A., Schwartz, E. N. F. & Schwartz, C. 1998. The largest lungless tetrapod: report on a second specimen of the lungless caecilian Atretochoana eiselti (Amphibia: Gymnophiona: Typhlonectidae) from Brazil. Journal of Natural History 32:617-627.

Wilkinson, M. & Thorley, J. L. 1998. Reduced supertrees. Trends in Evolution and Ecology 13:283.

Wilkinson, M., Upchurch, P., Barrett, P. M., Gower, D. J. & Benton, M. J. 1998. Robust dinosaur phylogeny Nature 396:423-424.

Wills, M. A., Briggs, D. E. G., Fortey, R. A., Wilkinson, M. & Sneath, P.H.A. 1998. An arthropod phylogeny based on fossil and recent taxa. Pp. 33-105. In (G. E. Edgecombe, ed.) Arthropod fossils and phylogeny. Colombia University Press.


Miller, P. J., Wilkinson, M. & Chen, I. 1997. Morphological and molecular phylogeny of gobioid fish (Teleostei: Gobioidei). Journal of Morphology 232:295.

Sheps, J. A., Wilkinson, M. & Orr, K. 1997. Novel cranial muscles and actions in a caecilian.Journal of Morphology 232:322.

Wilkinson, M. 1997. Limitations on the use of compatibility methods for polarizing and ordering characters. Systematic Biology 46:230-231.

Wilkinson, M. 1997. Congruence, consistency and the 'axioms' of cladistics. Taxon 46:739-742.

Wilkinson, M. 1997. Burning straw men sheds little light: a reply to Whiting and Kelly. Acta Biotheoretica, 45:17-27.

Wilkinson, M. 1997. Characters, congruence and quality: a study of neuroanatomical and traditional data in caecilian phylogeny. Biological Reviews 72:423-470.

Wilkinson, M. 1997. On phylogenetic relationships within Dendrotriton (Amphibia: Caudata: Plethodontidae): is there sufficient evidence? The Herpetological Journal 7:55-65.

Wilkinson, M. & Nussbaum, R. A. 1997. Comparative morphology and evolution of the lungless caecilian Atretochoana eiselti (Taylor) (Amphibia: Gymnophiona: Typhlonectidae). The Biological Journal of the Linnean Society 62:39-109.

Wilkinson, M., Thorley, J. L. & Benton, M. J. 1997. Uncertain turtle relationships. Nature 387:466.


Bruce, D. & Wilkinson M. 1996. Preliminary report on the amphibians of the Iwokrama rainforest. Glasgow University Expedition Society, Guyana '95 Expedition Report. Pp. 35-38.

Gower, D. J. & Wilkinson, M. 1996. Is there any consensus on basal archosaur phylogeny? Proceedings of the Royal Society B 263:1399-1406.

Wilkinson, M. 1996. Majority-rule reduced consensus methods and their use in bootstrapping. Molecular Biology and Evolution 13: 437-444.

Wilkinson, M. 1996. On the distribution of homoplasy and choosing among trees. Taxon 45:263-266.

Wilkinson, M. 1996. The taxonomic status of Typhlonectes venezuelense Fuhrmann (Amphibia: Gymnophiona: Typhlonectidae). The Herpetological Journal 6:30-31.

Wilkinson, M. 1996. The heart and aortic arches of rhinatrematid caecilians (Amphibia: Gymnophiona). The Zoological Journal of the Linnean Society 118:135-150.

Wilkinson, M. 1996. Resolution of the taxonomic status of Nectocaecilia haydee (Roze) and a key to the genera of the Typhlonectidae (Amphibia: Gymnophiona). Journal of Herpetology 30:413-415.

Wilkinson, M. & Benton, M. J. 1996. Sphenodontid phylogeny and the problems of multiple trees. Philosophical Transactions of the Royal Society B 351:1-16.

Wilkinson, M. & Bruce, D. 1996.< Preliminary report on the reptiles of the Iwokrama rainforest. Glasgow University Expedition Society, Guyana '95 Expedition Report.Pp. 41-43.

Wilkinson, M. & Nussbaum, R. A. 1996. On the phylogenetic position of the Uraeotyphlidae (Amphibia: Gymnophiona). Copeia 1996:550-562.

Wilkinson, M., Suter, S. J. & Shires, V. L. 1996. The reduced cladistic consensus method and cassiduloid echinoid phylogeny. Historical Biology 12:63-73.


Embley, T. M., Finlay, B. J., Dyal, P. L., Hirt, R. P., Wilkinson, M. & Williams, A. G. 1995. Multiple origins of anaerobic ciliates with hydrogenosomes within the radiation of aerobic ciliates. Proceedings of the Royal Society B 262:87-93.

Hirt, R. P., Dyal, P. L., Wilkinson, M., Finlay, B. J., Roberts, D. McL. & Embley, T. M. 1995. Phylogenetic relationships among karyolectids and heterotrichs inferred from small subunit rRNA sequences: resolution at the base of the ciliate tree. Molecular Phylogenetics and Evolution 4:77-87.

McInernay, J. O, Wilkinson, M., Patching, J. W., Embley, T. M. & Powell, R. 1995. Recovery and phylogenetic analysis of novel archeal ribosomal RNA sequences from a deep-sea deposit feeder. Applied and Environmental Microbiology 61:1646-1648.

Nussbaum, R. A. & Wilkinson, M. 1995. A new genus of lungless tetrapod: a radically divergent caecilian (Amphibia: Gymnophiona). Proceedings of the Royal Society B 261:331-335.

Wilkinson, M. 1995. A comparison of two methods of character construction. Cladistics 11:297-308.

Wilkinson, M.1995. Coping with missing entries in phylogenetic inference using parsimony. Systematic Biology 44:501-514.

Wilkinson, M.1995. Arbitrary resolutions, missing entries and the problem of zero-length branches in parsimony analysis. Systematic Biology 44:108-111.

Wilkinson, M. 1995. More on reduced consensus methods. Systematic Biology 44:435-439.

Wilkinson, M. & Benton, M. J. 1995. Missing data and rhynchosaur phylogeny. Historical Biology 10:137-150.

Wills, M. A., Briggs, D. E. G., Fortey, R. A. & Wilkinson, M. 1995. The significance of fossils in understanding arthropod evolution. Verhandlungen der Deutschen Zoologischen Gesellschaft 88.2:203-215.


Wilkinson, M. 1994. Weights and ranks in numerical phylogenetics Cladistics 10:321-329.

Wilkinson, M. 1994. Three-taxon statements: when is a parsimony analysis also a clique analysis? Cladistics 10:221-223.

Wilkinson, M. 1994. Common cladistic information and its consensus representation: reduced Adams and reduced cladistic consensus trees and profiles. Systematic Biology 43:343-368.

Wilkinson, M. 1994. The permutation method and character compatibility. Systematic Biology 43:274-277.


Wilkinson, M. 1992. Ordered versus unordered characters. Cladistics 8:375-385.

Wilkinson, M. 1992. On the life history of the caecilian genus Uraeotyphlus (Amphibia: Gymnophiona). Herpetological Journal 2:121-124.

Wilkinson, M. 1992. Novel modification of the tetrapod cardiovascular system in the West African Caecilian Herpele squalostoma (Amphibia: Gymnophiona: Caeciliaidae). Journal of Zoology 228:277-286.

Wilkinson, M. 1992. The phylogenetic position of the Rhinatrematidae (Amphibia: Gymnophiona): evidence from the larval lateral line system. Amphibia-Reptilia 13: 74-79.

Wilkinson, M. & Little, C. 1992. Review of Progressive Palaeontology. Palaeontology Newsletter 15:15

Wilkinson, M. & Nussbaum, R. A. 1992: Taxonomic status of Pseudosiphonops ptychodermis Taylor and Mimosiphonops vermiculatus Taylor (Amphibia: Gymnophiona: Caeciliaidae), with description of a new species. Journal of Natural History 26:675-688.


Wilkinson, M. 1991. The use of primitive character state distributions in the assessment of holophyly. Acta Biotheoretica 39:37-46.

Wilkinson, M. 1991. Homoplasy and parsimony analysis. Systematic Zoology 40:107-109.

Wilkinson, M. 1991. Adult tooth crown morphology in the Typhlonectidae (Amphibia: Gymnophiona): a reinterpretation of variation and its significance. Zeitschrift fur zoologische Systematische und Evolutions-forschung 29:304-311.


Wilkinson, M. 1990. A commentary on Ridley's cladistic solution to the species problem. Biology and Philosophy 5:433-446.

Wilkinson, M. 1990. The presence of a musculus retractor cloacae in female caecilians. Amphibia-Reptilia 11:300-304.


Nussbaum, R. A. & Wilkinson, M. 1989. On the classification and phylogeny of caecilians (Amphibia: Gymnophiona), a critical review. Herpetological Monographs 3:1-42.

Wilkinson, M. 1989. On the status of Nectocaecilia fasciata Taylor, with a discussion of the phylogeny of the Typhlonectidae (Amphibia: Gymnophiona). Herpetologica 45:23-36.


Wilkinson, M. 1988. Classical and evolutionary concepts of homology: a reply to Aboitiz. Acta Biotheoretica 37:315-319.

Wilkinson, M. 1988. The status of Nectocaecilia cooperi Taylor, with comments on the genus Nectocaecilia Taylor (Amphibia: Gymnophiona). Journal of Herpetology 22:119-121.

Wilkinson, M. & Nussbaum, R. A. 1988. Ichthyophiidae Taylor, 1968 (Amphibia: Gymnophiona): proposed conservation. Bulletin of Zoological Nomenclature 45:207-209.

Wilkinson, M. & Nussbaum, R. A. 1988. On the type locality of Chthonerpeton corrugatum Taylor (Amphibia: Gymnophiona). Herpetological Journal 1: 245-246.


Nussbaum, R. A. & Wilkinson, M. 1987. Two new species of Chthonerpeton (Amphibia: Gymnophiona: Typhlonectidae) from Brazil. Occasional Papers of the Museum of Zoology, University of Michigan 716: 1-15.

Wilkinson, M. 1983. An introduction to the caecilians. Herptile 8:43-47.

Wilkinson, M. 1981. Notes on a caecilian. Nectocaecilia sp. Herptile 5:22-25.


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